RRC ID 83631
Author Men Y, Hirayama S, Ao S, Sakurai Y, Shibata Y, Lo M, Sato Y, Murata S.
Title ESCRT-I and PTPN23 mediate microautophagy of ubiquitylated tau aggregates.
Journal J Cell Biol
Abstract Protein aggregates are degraded by both the autophagy-lysosomal and the ubiquitin-proteasome pathways. Macroautophagy and microautophagy, two forms of the autophagy-lysosomal pathway, are widely conserved across eukaryotes. While macroautophagy has been extensively studied in the context of degradation of protein aggregates, microautophagy remains less explored. Here, we identify the UBAP1-containing ESCRT-I complex and PTPN23 as new regulators for degradation of aggregated proteins through an unbiased genome-wide CRISPR knockout screen, using a cell line expressing tau repeat domain (tauRD) aggregates. ESCRT-I recognizes ubiquitylated tauRD via the UEV domain of TSG101. The accessory protein PTPN23, instead of ESCRT-II, bridges ESCRT-I and ESCRT-III to complete the endosomal microautophagy of ubiquitylated tauRD aggregates. Our results uncover the molecular mechanism underlying the degradation of tau aggregates by endosomal microautophagy.
Volume 224(6)
Published 2025-6-2
DOI 10.1083/jcb.202406120
PII 277371
PMID 40197510
PMC PMC11977513
MeSH Autophagy* DNA-Binding Proteins / genetics DNA-Binding Proteins / metabolism Endosomal Sorting Complexes Required for Transport* / genetics Endosomal Sorting Complexes Required for Transport* / metabolism Endosomes / metabolism HEK293 Cells HeLa Cells Humans Lysosomes / metabolism Microautophagy* Protein Aggregates Protein Tyrosine Phosphatases, Non-Receptor Proteolysis Transcription Factors / genetics Transcription Factors / metabolism Ubiquitination tau Proteins* / genetics tau Proteins* / metabolism
IF 8.811
Resource
Human and Animal Cells 293T(RCB2202)
DNA material CSII-EF-MCS (RDB04378) pCAG-HIVgp (RDB04394) pCMV-VSV-G-RSV-Rev (RDB04393)